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During embryonic development, the brain first forms as a tube, the anterior end of which enlarges into three hollow swellings that form the brain, and the posterior of which develops into the spinal cord. Some parts of the brain have changed little during vertebrate evolutionary history. Click here to view an diagram of the brain, and here for a clickable map of the brain.

Parts of the brain as seen from the middle of the brain. Image from Purves et al., Life: The Science of Biology, 4th Edition, by Sinauer Associates (www.sinauer.com) and WH Freeman (www.whfreeman.com), used with permission.

Vertebrate evolutionary trends include

  1. Increase in brain size relative to body size.
  2. Subdivision and increasing specialization of the forebrain, midbrain, and hindbrain.
  3. Growth in relative size of the forebrain, especially the cerebrum, which is associated with increasingly complex behavior in mammals.

The Brain Stem and Midbrain

The brain stem is the smallest and from an evolutionary viewpoint, the oldest and most primitive part of the brain. The brain stem is continuous with the spinal cord, and is composed of the parts of the hindbrain and midbrain. The medulla oblongata and pons control heart rate, constriction of blood vessels, digestion and respiration.

The midbrain consists of connections between the hindbrain and forebrain. Mammals use this part of the brain only for eye reflexes.

The Cerebellum

The cerebellum is the third part of the hindbrain, but it is not considered part of the brain stem. Functions of the cerebellum include fine motor coordination and body movement, posture, and balance. This region of the brain is enlarged in birds and controls muscle action needed for flight.

The Forebrain

The forebrain consists of the diencephalon and cerebrum. The thalamus and hypothalamus are the parts of the diencephalon. The thalamus acts as a switching center for nerve messages. The hypothalamus is a major homeostatic center having both nervous and endocrine functions.

The cerebrum, the largest part of the human brain, is divided into left and right hemispheres connected to each other by the corpus callosum. The hemispheres are covered by a thin layer of gray matter known as the cerebral cortex, the most recently evolved region of the vertebrate brain. Fish have no cerebral cortex, amphibians and reptiles have only rudiments of this area.

The cortex in each hemisphere of the cerebrum is between 1 and 4 mm thick. Folds divide the cortex into four lobes: occipital, temporal, parietal, and frontal. No region of the brain functions alone, although major functions of various parts of the lobes have been determined.

The major brain areas and lobes. Image from Purves et al., Life: The Science of Biology, 4th Edition, by Sinauer Associates (www.sinauer.com) and WH Freeman (www.whfreeman.com), used with permission.

The occipital lobe (back of the head) receives and processes visual information. The temporal lobe receives auditory signals, processing language and the meaning of words. The parietal lobe is associated with the sensory cortex and processes information about touch, taste, pressure, pain, and heat and cold. The frontal lobe conducts three functions:

  1. motor activity and integration of muscle activity
  2. speech
  3. thought processes

Functional areas of the brain. Image from Purves et al., Life: The Science of Biology, 4th Edition, by Sinauer Associates (www.sinauer.com) and WH Freeman (www.whfreeman.com), used with permission.

Most people who have been studied have their language and speech areas on the left hemisphere of their brain. Language comprehension is found in Wernicke's area. Speaking ability is in Broca's area. Damage to Broca's area causes speech impairment but not impairment of language comprehension. Lesions in Wernicke's area impairs ability to comprehend written and spoken words but not speech. The remaining parts of the cortex are associated with higher thought processes, planning, memory, personality and other human activities.

Parts of the cerebral cortex and the relative areas that are devoted to controlling various body regions. Image from Purves et al., Life: The Science of Biology, 4th Edition, by Sinauer Associates (www.sinauer.com) and WH Freeman (www.whfreeman.com), used with permission.


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