Themes > Science > Life Sciences > General Biology > Physiology > The Nervous System > Senses

Input to the nervous system is in the form of our five senses: pain, vision, taste, smell, and hearing. Vision, taste, smell, and hearing input are the special senses. Pain, temperature, and pressure are known as somatic senses. Sensory input begins with sensors that react to stimuli in the form of energy that is transmitted into an action potential and sent to the CNS.

Sensory Receptors

  • Sensory receptors are classified according to the type of energy they can detect and respond to.
  • Mechanoreceptors: hearing and balance, stretching.
  • Photoreceptors: light.
  • Chemoreceptors: smell and taste mainly, as well as internal sensors in the digestive and circulatory systems.
  • Thermoreceptors: changes in temperature.
  • Electroreceptors: detect electrical currents in the surrounding environment.

Mechanoreceptors vary greatly in the specific type of stimulus and duration of stimulus/action potentials. The most adaptable vertebrate mechanoreceptor is the hair cell. Hair cells are present in the lateral line of fish. In humans and mammals hair cells are involved with detection of sound and gravity and providing balance.

Hearing

Hearing involves the actions of the external ear, eardrum, ossicles, and cochlea. In hearing, sound waves in air are converted into vibrations of a liquid then into movement of hair cells in the cochlea. Finally they are converted into action potentials in a sensory dendrite connected to the auditory nerve. Very loud sounds can cause violent vibrations in the membrane under hair cells, causing a shearing or permanent distortion to the cells, resulting in permanent hearing loss.

Orientation and Gravity

Orientation and gravity are detected at the semicircular canals. Hair cells along three planes respond to shifts of liquid within the cochlea, providing a three-dimensional sense of equilibrium. Calcium carbonate crystals can shift in response to gravity, providing sensory information about gravity and acceleration.

Photoreceptors Detect Vision and Light Sensitivity

The human eye can detect light in the 400-700 nanometer (nm) range, a small portion of the electromagnetic spectrum, the visible light spectrum. Light with wavelengths shorter than 400 nm is termed ultraviolet (UV) light. Light with wavelengths longer than 700 nm is termed infrared (IR) light.

The electromagnetic spectrum. Image from Purves et al., Life: The Science of Biology, 4th Edition, by Sinauer Associates (www.sinauer.com) and WH Freeman (www.whfreeman.com), used with permission.

Eye

In the eye, two types of photoreceptor cells are clustered on the retina, or back portion of the eye. These receptors, rods and cones, apparently evolved from hair cells. Rods detect differences in light intensity; cones detect color. Rods are more common in a circular zone near the edge of the eye. Cones occur in the center (or fovea centralis) of the retina.

Light reaching a photoreceptor causes the breakdown of the chemical rhodopsin, which in turn causes a membrane potential that is transmitted to an action potential. The action potential transfers to synapsed neurons that connect to the optic nerve. The optic nerve connects to the occipital lobe of the brain.

Humans have three types of cones, each sensitive to a different color of light: red, blue and green. Opsins are chemicals that bind to cone cells and make those cells sensitive to light of a particular wavelength (or color). Humans have three different form of opsins coded for by three genes on the X chromosome. Defects in one or more of these opsin genes can cause color blindness, usually in males.


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